Williamston River Shingle

Site of Special Scientific Interest · 54.8627, -2.4984

Condition (Natural England)

Unfavourable Declining (1)
UnitFeatureConditionArea (ha)Assessed
NY681521 Unfavourable Declining 1.31 2011-11-10

Climate exposure

This is about exposure to a nationwide pressure this habitat type carries — not a verdict on this site's own management.

Coastal

Coastal habitats are exposed to sea-level rise and storm erosion. Where hard sea defences sit behind them, there's nowhere landward to retreat to — a squeeze that keeps compressing the habitat no matter how well it's managed.

Geology & landscape

Mining / Industrial: Mine Spoil · Rock Type: Limestone

Landscape processes: Coastal

Description

Area, 1.5 ha. 3.7 ac. Description Williamston River Shingle situated on the South Tyne, is one of a series of metalliferous shingle sites, a habitat which is restricted in Northumberland to the South Tyne, West and East Allen rivers. The shingle contains high levels of heavy metals and supports an unusual community of metal tolerant plants. The site also shows the succession from bare shingle and grassland to scrub and finally woodland. The Pennines in south Northumberland were worked for their rich seams of lead and zinc until the early part of this century. The extraction of the metals by washing and sedimentation processes allowed the release of fine fractions of lead, zinc, barium and cadmium into the rivers. During flooding these metals were-deposited with sand in some of the shingle beds along the rivers. Although heavy metals are toxic to most plants, certain tolerant species are able to grow on the shingles. Of these spring sandwort Minuartia verna and alpine pennycress Noccaea caerulescens are normally found in limestone grassland in upland areas, commonly associated with mine spoil; mountain pansy Viola lutea is also usually restricted to upland areas, whilst common scurvy grass Cochlearia officinalis and thrift Armeria maritima are normally found on the coast. The shingle also supports a rich community of metal tolerant lichens. Other plants of the short, dry grassland on the shingle are red fescue Festuca rubra, common thyme Thymus praecox, quaking grass Briza media, lady’s bedstraw Galium verum and kidney vetch Anthyllis vulneraria. Areas of tall grassland are dominated by downy oat grass Avenula pubescens, false oat grass Arrhenatherum elatius and lesser knapweed Centaurea nigra. Where the sandy soil is rich in lime the flora is very diverse, with plants such as meadow oat grass Avenula pratensis, purging flax Linum catharticum salad burnet Poterium sanguisorba and yellow oat grass Trisetum flavescens. Scrub, comprising gorse Ulex europaeus, downy birch Betula pubescens, goat willow Salix caprea and alder Alnus glutinosa is present along much of the riverbank and is encroaching on the grassland in places. It has developed into secondary birch woodland where long established and has a grassy field layer of Yorkshire fog Holcus lanatus, common bent Agrostis capillaris, wood meadow grass Poa nemoralis and pignut Conopodium majus. The woodland supports a large population of narrow lipped helleborine Epipactis leptochila, an uncommon plant which has its only known localities in northern Britain on the heavy metal shingles of Northumberland. At the north end of the site alder and ash Fraxinus excelsior occur along the river bank with a field layer which includes red campion Silene dioica, dog’s mercury Mercurialis perennis and bluebell Hyacinthoides non-scripta. Other Information This site is managed as a nature reserve by the Northumberland Wildlife Trust.

Species mentioned

English name
search for this in other sites
Latin name
search with Google
IUCN / Red List Type Family
common bent Agrostis capillaris LC grass poaceae
false oat grass Arrhenatherum elatius LC grass poaceae
meadow oat grass Avenula pratensis Not listed in GB list grass poaceae
downy oat grass Avenula pubescens LC grass poaceae
quaking grass Briza media VU grass poaceae
red fescue Festuca rubra LC grass poaceae
Yorkshire fog Holcus lanatus LC grass poaceae
wood meadow grass Poa nemoralis LC grass poaceae
yellow oat grass Trisetum flavescens LC grass poaceae
lady's fingers (kidney vetch) Anthyllis vulneraria NT herb fabaceae
thrift Armeria maritima CR herb plumbaginaceae
black knapweed Centaurea nigra LC herb asteraceae
scurvy grass Cochlearia officinalis LC herb brassicaceae
pignut Conopodium majus LC herb apiaceae
narrow-lipped helleborine Epipactis leptochila VU herb orchidaceae
lady's bedstraw Galium verum LC herb rubiaceae
bluebell Hyacinthoides non-scripta LC herb liliaceae
fairy flax Linum catharticum LC herb linaceae
dog's mercury Mercurialis perennis LC herb euphorbiaceae
spring sandwort Minuartia verna Not listed in GB list herb carophyllaceae
alpine pennycress Noccaea caerulescens VU herb brassicaceae
salad burnet Poterium sanguisorba VU herb rosaceae
red campion Silene dioica LC herb carophyllaceae
creeping thyme Thymus praecox Not listed in GB list herb lamiaceae
yellow vetch Viola lutea LC herb violaceae
gorse Ulex europaeus LC shrub leguminosae
alder Alnus glutinosa LC tree salicaceae
downy birch Betula pubescens LC tree betulaceae
ash Fraxinus excelsior LC tree oleaceae
goat willow Salix caprea LC tree shrub salicaceae

Insects to look out for

These moths, butterflies and beetles are known to feed on plants recorded at this site, and carry a notable UK conservation status — but a host-plant being present doesn't mean the insect has actually been recorded here. Treat this as "worth watching for," not a confirmed sighting.

SpeciesStatusVia host-plant
Black-veined moth
Siona lineata
Critically Endangered Centaurea nigra
Straw belle
Aspitates gilvaria
Endangered Linum catharticum
The lappet
Gastropacha quercifolia
Endangered Salix spp.
Scarce vapourer
Orgyia recens
Endangered Alnus glutinosa, Betula spp., Salix spp.
Mouse moth
Amphipyra tragopoginis
Vulnerable Poterium sanguisorba, Salix caprea
The sprawler
Asteroscopus sphinx
Vulnerable Betula spp., Fraxinus excelsior, Salix spp.
Broom moth
Ceramica pisi
Vulnerable Betula spp.
Scarce chocolate-tip
Clostera anachoreta
Vulnerable Salix spp.
Mottled umber
Erannis defoliaria
Vulnerable Betula spp., Salix spp.
Satyr pug
Eupithecia satyrata
Vulnerable Centaurea spp., Galium spp., Salix spp.
White-line dart
Euxoa tritici
Vulnerable Galium verum
Grayling
Hipparchia semele
Vulnerable Festuca rubra
Bedstraw hawk-moth
Hyles gallii
Vulnerable Galium spp.
The lackey
Malacosoma neustria
Vulnerable Salix spp.
Pale brindled
Phigalia pilosaria
Vulnerable Alnus glutinosa, Betula pubescens, Fraxinus excelsior, Salix caprea
Silver studded blue
Plebejus argus
Vulnerable Ulex spp.
Grizzled skipper
Pyrgus malvae
Vulnerable Poterium sanguisorba
Lead belle
Scotopteryx mucronata
Vulnerable Ulex europaeus
Pale eggar
Trichiura crataegi
Vulnerable Betula spp., Salix spp.
Flounced chestnut
Agrochola helvola
Near Threatened Betula spp., Salix spp.
Beaded chestnut
Agrochola lychnidis
Near Threatened Salix spp.
Grey chi
Antitype chi
Near Threatened Armeria maritima, Salix caprea
Purple emperor
Apatura iris
Near Threatened Salix caprea
Minor shoulder-knot
Brachylomia viminalis
Near Threatened Salix caprea
The sallow
Cirrhia icteritia
Near Threatened Salix spp.
Small heath
Coenonympha pamphilus
Near Threatened Agrostis spp., Festuca spp., Poa spp.
Small blue
Cupido minimus
Near Threatened Anthyllis vulneraria
Dingy mocha
Cyclophora pendularia
Near Threatened Alnus glutinosa, Salix caprea
Kentish glory
Endromis versicolora
Near Threatened Alnus glutinosa, Betula pubescens
Large thorn
Ennomos autumnaria
Near Threatened Alnus glutinosa, Betula spp., Salix spp.
September thorn
Ennomos erosaria
Near Threatened Betula spp.
Dusky thorn
Ennomos fuscantaria
Near Threatened Fraxinus excelsior
Autumnal rustic
Eugnorisma glareosa
Near Threatened Betula spp., Galium spp., Hyacinthoides non-scripta
Grey pug
Eupithecia subfuscata
Near Threatened Centaurea spp., Salix spp.
Double dart
Graphiphora augur
Near Threatened Betula spp., Salix spp.
Scarce forester
Jordanita globulariae
Near Threatened Centaurea nigra
Wall
Lasiommata megera
Near Threatened Agrostis spp., Holcus lanatus
Rosy minor
Litoligia literosa
Near Threatened Arrhenatherum elatius
Dark-barred twin-spot carpet
Xanthorhoe ferrugata
Near Threatened Galium spp.
White prominent
Leucodonta bicoloria
Regionally Extinct Betula spp.

Host-plant data: Cook et al. (2025), Traits data for the butterflies and macro-moths of Great Britain and Ireland, 2024, UK Centre for Ecology & Hydrology / Butterfly Conservation; and Padovani et al. (2020), Database of Insects and their Food Plants (DBIF), UK Centre for Ecology & Hydrology (both Open Government Licence). Contains data supplied by UK Centre for Ecology & Hydrology.

Plant communities

OV37 — Calaminarian grassland

Festuca ovina-Minuartia verna community

Matches 10 species from the description above: Agrostis capillaris, Anthyllis vulneraria, Arrhenatherum elatius, Avenula pratensis, Festuca rubra, Linum catharticum, Minuartia verna, Noccaea caerulescens, Thymus praecox, Viola lutea

Calluna vulgaris, Carex caryophyllea, Carex flacca, Bryum pallens, Pohlia nutans, Dicranum scoparium, Rhytidiadelphus squarrosus, Weissia controversa, Cladonia arbuscula, Cladonia chlorophaea, Cladonia furcata, Cladonia gracilis, Cladonia portentosa, Cladonia pyxidata, Cladonia rangiformis, Coelocaulon aculeatum, Parmelia saxatilis, Achillea millefolium, Cirsium vulgare, Jacobaea vulgaris, Pilosella officinarum, Noccaea caerulescens, Campanula rotundifolia, Cerastium fontanum, Minuartia verna, Anthyllis vulneraria, Lotus corniculatus, Trifolium repens, Geranium robertianum, Thymus praecox, Linum catharticum, Euphrasia officinalis, Plantago lanceolata, Rumex acetosa, Galium saxatile, Galium sterneri, Viola lutea, Agrostis capillaris, Agrostis stolonifera, Anthoxanthum odoratum, Arrhenatherum elatius, Avenula pratensis, Dactylis glomerata, Festuca ovina, Festuca rubra, Koeleria macrantha

W22 — Hedgerow scrub, Mixed scrub

Prunus spinosa-Rubus fruticosus scrub

Matches 8 species from the description above: Agrostis capillaris, Armeria maritima, Festuca rubra, Holcus lanatus, Hyacinthoides non-scripta, Mercurialis perennis, Silene dioica, Ulex europaeus

Hedera helix, Acer campestre, Corylus avellana, Ulex europaeus, Prunus spinosa, Plagiomnium undulatum, Brachythecium rutabulum, Eurhynchium striatum, Kindbergia praelonga, Thuidium tamariscinum, Atrichum undulatum, Lophocolea bidentata, Achillea millefolium, Cardamine flexuosa, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Silene uniflora, Stellaria media agg., Mercurialis perennis, Geranium robertianum, Stachys sylvatica, Teucrium scorodonia, Hyacinthoides non-scripta, Oxalis acetosella, Digitalis purpurea, Plantago lanceolata, Veronica chamaedrys, Armeria maritima, Rumex acetosa, Filipendula ulmaria, Geum urbanum, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Urtica dioica, Viola riviniana, Agrostis capillaris, Brachypodium sylvaticum, Dactylis glomerata, Festuca rubra, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris borreri, Dryopteris filix-mas

W21 — Hedgerow scrub, Mixed scrub

Crataegus monogyna-Hedera helix scrub

Matches 7 species from the description above: Arrhenatherum elatius, Fraxinus excelsior, Holcus lanatus, Hyacinthoides non-scripta, Mercurialis perennis, Poterium sanguisorba, Silene dioica

Hedera helix, Salix cinerea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Corylus avellana, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Malus sylvestris, Sorbus aria, Taxus baccata, Ulmus glabra, Ulmus procera, Sambucus nigra, Viburnum lantana, Euonymus europaeus, Cornus sanguinea, Juniperus communis, Ligustrum vulgare, Clematis vitalba, Rhamnus cathartica, Crataegus laevigata, Crataegus monogyna, Prunus spinosa, Juncus effusus, Plagiomnium undulatum, Brachythecium rutabulum, Kindbergia praelonga, Rhynchostegium confertum, Tamus communis, Heracleum sphondylium, Torilis japonica, Arum maculatum, Arctium minus, Cirsium arvense, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Stellaria graminea, Stellaria holostea, Stellaria media agg., Calystegia sepium, Mercurialis perennis, Geranium robertianum, Iris foetidissima, Glechoma hederacea, Lamium purpureum, Origanum vulgare, Stachys sylvatica, Teucrium scorodonia, Allium ursinum, Hyacinthoides non-scripta, Circaea lutetiana, Rumex sanguineus, Anemone nemorosa, Ranunculus repens, Fragaria vesca, Geum urbanum, Potentilla sterilis, Poterium sanguisorba, Rosa arvensis, Rosa canina, Rubia peregrina, Rubus fruticosus agg., Galium aparine, Urtica dioica, Viola riviniana, Anisantha sterilis, Arrhenatherum elatius, Brachypodium pinnatum, Brachypodium sylvaticum, Bromopsis erecta, Dactylis glomerata, Elytrigia repens, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris filix-mas

CG12 — Alpine and subalpine calcareous grassland, High montane heath and snow bed

Festuca ovina-Alchemilla alpina-Silene acaulis dwarf-herb community

Matches 6 species from the description above: Agrostis capillaris, Armeria maritima, Festuca rubra, Linum catharticum, Thymus praecox, Viola lutea

Empetrum nigrum, Calluna vulgaris, Kalmia procumbens, Vaccinium myrtillus, Salix herbacea, Carex bigelowii, Carex demissa, Carex panicea, Carex pilulifera, Carex pulicaris, Juncus trifidus, Juncus triglumis, Luzula campestris, Luzula multiflora, Luzula spicata, Luzula sylvatica, Diphasiastrum alpinum, Huperzia selago, Selaginella selaginoides, Andreaea alpina, Aulacomnium turgidum, Bartramia ithyphylla, Breutelia chrysocoma, Mnium hornum, Dicranum scoparium, Ditrichum flexicaule, Racomitrium canescens sens.lat., Racomitrium fasciculare, Racomitrium lanuginosum, Ctenidium molluscum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Hypnum hamulosum, Lescuraea incurvata, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Thuidium delicatulum, Thuidium tamariscinum, Pogonatum urnigerum, Polytrichastrum alpinum, Tortella tortuosa, Barbilophozia hatcheri, Barbilophozia lycopodioides, Diplophyllum albicans, Frullania tamarisci, Nardia scalaris, Plagiochila asplenioides, Ptilidium ciliare, Scapania subalpina, Tritomaria quinquedentata, Cetraria islandica, Cladonia arbuscula, Cladonia gracilis, Cladonia pyxidata, Cladonia uncialis, Peltigera canina, Achillea millefolium, Antennaria dioica, Gnaphalium supinum, Hieraclium spp., Scorzoneroides autumnalis, Solidago virgaurea, Taraxacum officianalis, Myosotis alpestris, Cardamine pratensis, Draba norvegica, Campanula rotundifolia, Succisa pratensis, Cerastium alpinum, Cerastium fontanum, Minuartia sedoides, Sagina saginoides, Silene acaulis, Sedum rosea, Vaccinium vitis-idaea, Thymus praecox, Linum catharticum, Epilobium anagallidifolium, Euphrasia frigida, Euphrasia officinalis, Oxalis acetosella, Plantago maritima, Veronica alpina, Veronica officinalis, Veronica serpyllifolia, Armeria maritima, Polygonum viviparum, Rumex acetosa, Anemone nemorosa, Ranunculus acris, Thalictrum alpinum, Trollius europaeus, Alchemilla alpina, Alchemilla filicaulis, Filipendula ulmaria, Geum rivale, Potentilla crantzii, Potentilla erecta, Sibbaldia procumbens, Galium boreale, Galium saxatile, Saxifraga aizoides, Saxifraga hypnoides, Saxifraga nivalis, Saxifraga oppositifolia, Saxifraga stellaris, Viola lutea, Viola palustris, Viola riviniana, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Festuca vivipara, Nardus stricta, Poa pratensis, Botrychium lunaria

SD3 — Coastal vegetated shingle, Sand dune

Matricaria maritima-Galium aparine strandline community

Matches 4 species from the description above: Armeria maritima, Cochlearia officinalis, Festuca rubra, Holcus lanatus

Bolboschoenus maritimus, Juncus bufonius, Juncus gerardii, Ligusticum scoticum, Matricaria discoidea, Sonchus arvensis, Sonchus asper, Tripleurospermum maritimum, Mertensia maritima, Cakile maritima, Cochlearia officinalis, Honckenya peploides, Stellaria media agg., Atriplex glabriuscula, Atriplex patula, Atriplex prostrata, Triglochin maritimum, Plantago maritima, Armeria maritima, Rumex crispus, Glaux maritima, Galium aparine, Agrostis stolonifera, Elytrigia repens, Festuca rubra, Holcus lanatus, Leymus arenarius, Poa annua

U13 — High montane heath and snow bed, Upland dry heath

Deschampsia cespitosa-Galium saxatile grassland

Matches 4 species from the description above: Agrostis capillaris, Armeria maritima, Festuca rubra, Thymus praecox

Vaccinium myrtillus, Salix herbacea, Carex bigelowii, Carex pilulifera, Luzula multiflora, Luzula sylvatica, Diphasiastrum alpinum, Huperzia selago, Selaginella selaginoides, Pohlia nutans, Kiaeria starkei, Racomitrium lanuginosum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Plagiothecium undulatum, Pleurozium schreberi, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Polytrichastrum alpinum, Polytrichum commune, Sphagnum capillifolium, Sphagnum russowii, Anastrepta orcadensis, Barbilophozia floerkei, Barbilophozia lycopodioides, Nardia scalaris, Ptilidium ciliare, Cetraria islandica, Cladonia gracilis, Peltigera canina, Achillea millefolium, Gnaphalium supinum, Saussurea alpina, Campanula rotundifolia, Cerastium alpinum, Cerastium fontanum, Minuartia sedoides, Silene acaulis, Vaccinium vitis-idaea, Thymus praecox, Euphrasia frigida, Oxalis acetosella, Armeria maritima, Polygonum viviparum, Rumex acetosa, Ranunculus acris, Alchemilla alpina, Alchemilla filicaulis, Alchemilla glabra, Potentilla erecta, Sibbaldia procumbens, Galium saxatile, Saxifraga hypnoides, Saxifraga stellaris, Viola palustris, Viola riviniana, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Nardus stricta, Blechnum spicant, Dryopteris dilatata

U14 — High montane heath and snow bed, Upland dry heath

Alchemilla alpina-Sibbaldia procumbens dwarf-herb community

Matches 4 species from the description above: Agrostis capillaris, Armeria maritima, Festuca rubra, Thymus praecox

Vaccinium myrtillus, Salix herbacea, Carex bigelowii, Carex pilulifera, Juncus trifidus, Luzula multiflora, Luzula spicata, Diphasiastrum alpinum, Huperzia selago, Selaginella selaginoides, Andreaea alpina, Pohlia nutans, Dicranum scoparium, Racomitrium aquaticum, Racomitrium canescens sens.lat., Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Ctenidium molluscum, Hylocomium splendens, Hypnum callichroum, Pleurozium schreberi, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Oligotrichum hercynicum, Pogonatum urnigerum, Polytrichastrum alpinum, Polytrichum piliferum, Barbilophozia floerkei, Diplophyllum albicans, Lophozia sudetica, Nardia scalaris, Ptilidium ciliare, Cetraria islandica, Cladonia arbuscula, Cladonia bellidiflora, Cladonia pyxidata, Achillea millefolium, Gnaphalium supinum, Scorzoneroides autumnalis, Campanula rotundifolia, Cerastium alpinum, Cerastium fontanum, Minuartia sedoides, Sagina saginoides, Silene acaulis, Thymus praecox, Euphrasia officinalis, Armeria maritima, Polygonum viviparum, Rumex acetosa, Ranunculus acris, Thalictrum alpinum, Trollius europaeus, Alchemilla alpina, Alchemilla filicaulis, Alchemilla glabra, Potentilla erecta, Sibbaldia procumbens, Galium saxatile, Saxifraga hypnoides, Saxifraga oppositifolia, Saxifraga stellaris, Viola palustris, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Nardus stricta, Blechnum spicant, Botrychium lunaria

MC6 — Coastal vegetated shingle, Hard cliff and cliff slope

Atriplex prostrata-Beta vulgaris ssp. maritima sea-bird cliff community

Matches 3 species from the description above: Armeria maritima, Cochlearia officinalis, Festuca rubra

Daucus carota, Sonchus oleraceus, Taraxacum officianalis, Tripleurospermum maritimum, Cochlearia officinalis, Silene uniflora, Spergularia rupicola, Atriplex hastata, Malva arborea, Plantago coronopus, Armeria maritima, Polygonum aviculare, Rumex crispus, Bromus hordeaceous, Catapodium marinum, Dactylis glomerata, Festuca rubra

SD1 — Coastal vegetated shingle, Sand dune

Rumex crispus-Glaucium flavum shingle community

Matches 3 species from the description above: Arrhenatherum elatius, Festuca rubra, Holcus lanatus

Solanum dulcamara, Crithmum maritimum, Cirsium arvense, Cirsium vulgare, Helminthotheca echioides, Hypochaeris radicata, Jacobaea vulgaris, Lactuca serriola, Leontodon saxatilis, Senecio viscosus, Sonchus arvensis, Sonchus asper, Tripleurospermum maritimum, Crambe maritima, Cerastium fontanum, Honckenya peploides, Sagina apetala, Silene uniflora, Atriplex prostrata, Sedum acre, Euphorbia paralias, Lathyrus japonicus, Geranium robertianum, Glaucium flavum, Plantago coronopus, Plantago lanceolata, Rumex crispus, Potentilla anserina, Ammophila arenaria, Arrhenatherum elatius, Elytrigia atherica, Elytrigia juncea, Festuca rubra, Holcus lanatus, Lolium perenne

U10 — High montane heath and snow bed, Lowland dry acid grassland, Upland dry heath

Carex bigelowii-Racomitrium lanuginosum moss-heath

Matches 3 species from the description above: Agrostis capillaris, Armeria maritima, Thymus praecox

Empetrum nigrum, Kalmia procumbens, Vaccinium myrtillus, Salix herbacea, Carex bigelowii, Carex panicea, Carex pilulifera, Juncus squarrosus, Juncus trifidus, Luzula spicata, Diphasiastrum alpinum, Huperzia selago, Selaginella selaginoides, Andreaea alpina, Aulacomnium turgidum, Pohlia nutans, Campylopus flexuosus, Dicranum fuscescens, Dicranum scoparium, Kiaeria starkei, Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Hypnum hamulosum, Pleurozium schreberi, Rhytidiadelphus loreus, Sanionia uncinata, Polytrichastrum alpinum, Polytrichum juniperinum, Polytrichum longisetum, Polytrichum piliferum, Polytrichum strictum, Anastrepta orcadensis, Diplophyllum albicans, Nardia scalaris, Ptilidium ciliare, Alectoria nigricans, Cetraria islandica, Cladonia arbuscula, Cladonia cervicornis, Cladonia coccifera, Cladonia crispata, Cladonia furcata, Cladonia gracilis, Cladonia impexa, Cladonia pyxidata, Cladonia rangiferina, Cladonia squamosa, Cladonia uncialis, Cornicularia aculeata, Ochrolechia frigida, Peltigera canina, Sphaerophorus globosus, Thamnolia vermicularis, Achillea millefolium, Antennaria dioica, Gnaphalium supinum, Solidago virgaurea, Campanula rotundifolia, Succisa pratensis, Minuartia sedoides, Silene acaulis, Vaccinium vitis-idaea, Thymus praecox, Armeria maritima, Polygonum viviparum, Rumex acetosa, Alchemilla alpina, Potentilla erecta, Sibbaldia procumbens, Galium saxatile, Viola palustris, Viola riviniana, Agrostis capillaris, Agrostis vinealis, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Nardus stricta

W20 — Montane scrub

Salix lapponum-Luzula sylvatica scrub

Matches 3 species from the description above: Agrostis capillaris, Festuca rubra, Thymus praecox

Salix arbuscula, Salix cinerea, Salix lapponum, Salix myrsinites, Salix phylicifolia, Salix reticulata, Empetrum nigrum, Calluna vulgaris, Salix lanata, Vaccinium myrtillus, Vaccinium uliginosum, Carex atrata, Carex bigelowii, Carex binervis, Carex flacca, Luzula multiflora, Luzula sylvatica, Huperzia selago, Selaginella selaginoides, Aulacomnium palustre, Mnium hornum, Philonotis fontana, Plagiomnium undulatum, Rhizomnium punctatum, Dicranum bonjeanii, Dicranum majus, Dicranum scoparium, Fissidens adianthoides, Racomitrium lanuginosum, Calliergonell cuspidatum, Ctenidium molluscum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Palustriella commutatum, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Ptilium crista-castrensis, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Scorpidium revolvens, Thuidium tamariscinum, Atrichum undulatum, Polytrichastrum alpinum, Sphagnum capillifolium, Sphagnum fallax, Sphagnum girgensohnii, Sphagnum palustre, Sphagnum russowii, Sphagnum squarrosum, Sphagnum subnitens, Sphagnum teres, Barbilophozia barbata, Chiloscyphus polyanthos, Diplophyllum albicans, Lophocolea bidentata, Plagiochila porelloides, Ptilidium ciliare, Scapania undulata, Cladonia squamosa, Peltigera canina, Angelica sylvestris, Caltha palustris, Hieraclium spp., Saussurea alpina, Solidago virgaurea, Campanula rotundifolia, Succisa pratensis, Valeriana officinalis, Rhodiola rosea, Vaccinium vitis-idaea, Thymus praecox, Epilobium anagallidifolium, Dactylorhiza viridis, Euphrasia officinalis, Rhinanthus minor, Oxalis acetosella, Oxyria digyna, Polygonum viviparum, Rumex acetosa, Anemone nemorosa, Ranunculus acris, Thalictrum alpinum, Alchemilla alpina, Alchemilla glabra, Filipendula ulmaria, Geum rivale, Galium boreale, Galium saxatile, Saxifraga aizoides, Saxifraga hypnoides, Saxifraga oppositifolia, Saxifraga stellaris, Viola riviniana, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Nardus stricta, Dryopteris dilatata, Phegopteris connectilis

SD2 — Coastal vegetated shingle, Sand dune

Honkenya peploides-Cakile maritima strandline community

Matches 2 species from the description above: Arrhenatherum elatius, Festuca rubra

Salsola kali, Achillea millefolium, Cirsium arvense, Eryngium maritimum, Hypochaeris radicata, Senecio vulgaris, Sonchus arvensis, Tripleurospermum maritimum, Cakile maritima, Cerastium diffusum, Honckenya peploides, Silene uniflora, Stellaria media agg., Atriplex glabriuscula, Atriplex laciniata, Atriplex littoralis, Atriplex patula, Atriplex prostrata, Sedum acre, Polygonum oxyspermum subsp. Raii, Rumex crispus, Potentilla anserina, Agrostis stolonifera, Ammophila arenaria, Arrhenatherum elatius, Elytrigia juncea, Elytrigia repens, Festuca rubra, Leymus arenarius, Poa annua

U7 — High montane heath and snow bed

Nardus stricta-Carex bigelowii grass-heath

Matches 2 species from the description above: Agrostis capillaris, Thymus praecox

Empetrum nigrum, Calluna vulgaris, Vaccinium myrtillus, Vaccinium uliginosum, Salix herbacea, Carex bigelowii, Carex nigra, Carex panicea, Carex pilulifera, Eriophorum vaginatum, Trichophorum cespitosum, Juncus squarrosus, Juncus trifidus, Luzula multiflora, Luzula sylvatica, Diphasiastrum alpinum, Huperzia selago, Campylopus flexuosus, Dicranum fuscescens, Dicranum scoparium, Kiaeria starkei, Racomitrium lanuginosum, Hylocomium splendens, Hypnum cupressiforme, Plagiothecium undulatum, Pleurozium schreberi, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Polytrichastrum alpinum, Polytrichum commune, Polytrichum juniperinum, Polytrichum piliferum, Polytrichum strictum, Sphagnum capillifolium, Anastrepta orcadensis, Anastrophyllum donianum, Barbilophozia floerkei, Barbilophozia hatcheri, Bazzania tricrenata, Diplophyllum albicans, Lophozia sudetica, Pleurozia purpurea, Ptilidium ciliare, Scapania gracilis, Scapania nemorea, Cetraria delisei, Cetraria islandica, Cladonia arbuscula, Cladonia bellidiflora, Cladonia coccifera, Cladonia gracilis, Cladonia impexa, Cladonia leucophaea, Cladonia pyxidata, Cladonia rangiferina, Cladonia tenuis, Cladonia uncialis, Gnaphalium supinum, Solidago virgaurea, Cornus suecica, Vaccinium vitis-idaea, Thymus praecox, Narthecium ossifragum, Rumex acetosa, Alchemilla alpina, Potentilla erecta, Galium saxatile, Viola palustris, Agrostis capillaris, Agrostis vinealis, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Nardus stricta

U9 — High montane heath and snow bed

Juncus trifidus-Racomitrium lanuginosum rush-heath

Matches 2 species from the description above: Agrostis capillaris, Armeria maritima

Empetrum nigrum, Kalmia procumbens, Vaccinium myrtillus, Vaccinium uliginosum, Salix herbacea, Carex bigelowii, Carex pilulifera, Juncus trifidus, Luzula spicata, Diphasiastrum alpinum, Huperzia selago, Dicranum fuscescens, Racomitrium lanuginosum, Pleurozium schreberi, Polytrichastrum alpinum, Polytrichum piliferum, Barbilophozia floerkei, Diplophyllum albicans, Ptilidium ciliare, Cetraria delisei, Cetraria islandica, Cetraria nivalis, Cladonia arbuscula, Cladonia bellidiflora, Cladonia coccifera, Cladonia crispata, Cladonia gracilis, Cladonia pyxidata, Cladonia rangiferina, Cladonia uncialis, Cornicularia aculeata, Ochrolechia frigida, Sphaerophorus globosus, Stereocaulon vesuvianum, Thamnolia vermicularis, Silene acaulis, Vaccinium vitis-idaea, Armeria maritima, Polygonum viviparum, Alchemilla alpina, Galium saxatile, Agrostis capillaris, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Nardus stricta, Nitella flexilis

U11 — High montane heath and snow bed, Lowland dry acid grassland, Upland dry heath

Polytrichum sexangulare-Kiaeria starkei snow-bed

Matches 1 species from the description above: Agrostis capillaris

Empetrum nigrum, Salix herbacea, Carex bigelowii, Juncus trifidus, Huperzia selago, Conostomum tetragonum, Pohlia drummondii, Pohlia ludwigii, Pohlia nutans, Dicranum fuscescens, Kiaeria falcata, Kiaeria starkei, Racomitrium canescens sens.lat., Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Pleurozium schreberi, Rhytidiadelphus loreus, Oligotrichum hercynicum, Polytrichastrum alpinum, Polytrichastrum sexangulare, Anthelia juratzkana, Barbilophozia floerkei, Lophozia sudetica, Marsupella emarginata, Moerckia blyttii, Nardia scalaris, Pleurocladula albescens, Cetraria islandica, Cladonia bellidiflora, Cladonia pyxidata, Cladonia uncialis, Solorina crocea, Gnaphalium supinum, Cerastium cerastoides, Silene acaulis, Alchemilla alpina, Sibbaldia procumbens, Galium saxatile, Saxifraga stellaris, Viola palustris, Agrostis capillaris, Deschampsia cespitosa, Deschampsia flexuosa, Festuca vivipara, Nardus stricta, Cryptogramma crispa, Blechnum spicant, Athyrium distentifolium

U8 — High montane heath and snow bed

Carex bigelowii-Polytrichum alpinum sedge-heath

Matches 1 species from the description above: Agrostis capillaris

Vaccinium myrtillus, Salix herbacea, Carex bigelowii, Juncus trifidus, Luzula spicata, Huperzia selago, Pohlia nutans, Dicranum fuscescens, Dicranum scoparium, Racomitrium canescens sens.lat., Racomitrium fasciculare, Racomitrium lanuginosum, Hylocomium splendens, Pleurozium schreberi, Sanionia uncinata, Polytrichastrum alpinum, Sphagnum compactum, Ptilidium ciliare, Cetraria delisei, Cetraria islandica, Cladonia arbuscula, Cladonia bellidiflora, Cladonia coccifera, Cladonia floerkeana, Cladonia gracilis, Cladonia macrophylla, Cladonia pyxidata, Cladonia rangiferina, Cladonia squamosa, Cladonia uncialis, Cornicularia aculeata, Ochrolechia frigida, Galium saxatile, Saxifraga stellaris, Agrostis capillaris, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Nardus stricta

A14 — Fast flowing river and stream

Myriophyllum alterniflorum community

Isolepis fluitans, Juncus bulbosus, Fontinalis antipyretica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Lemna minor, Callitriche stagnalis, Littorella uniflora, Potamogeton filiformis, Potamogeton gramineus, Potamogeton natans, Ranunculus peltatus, Chara spp., Nitella hyalina

A17 — Chalk river and stream, Fast flowing river and stream

Ranunculus penicillatus ssp. pseudofluitans community

Berula erecta, Nasturtium officinale, Elodea canadensis, Lemna minor, Callitriche stagnalis, Veronica beccabunga, Polygonum amphibium, Polygonum hydropiper, Potamogeton crispus, Potamogeton pectinatus, Ranunculus penicillatus ssp. pseudofluitans

A18 — Fast flowing river and stream

Ranunculus fluitans community

Fontinalis antipyretica, Fontinalis squamosa var. curnowii, Myriophyllum alterniflorum, Myriophyllum spicatum, Elodea canadensis, Mentha aquatica, Lemna gibba, Lemna minor, Potamogeton perfoliatus, Ranunculus fluitans, Glyceria fluitans, Phalaris arundinacea

A9 — Fast flowing river and stream, Marginal and inundation, Pond ditch and canal, Sluggish river and stream

Potamogeton natans community

Schoenoplectus lacustris, Juncus bulbosus, Sparganium angustifolium, Sparganium erectum, Sparganium minimum, Alisma plantago-aquatica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Elodea canadensis, Lemna minor, Lemna trisulca, Nymphaea alba, Callitriche platycarpa, Callitriche stagnalis, Littorella uniflora, Potamogeton natans, Ranunculus trichophyllus, Glyceria fluitans, Chara subspinosa

U12 — High montane heath and snow bed, Upland dry heath

Salix herbacea-Racomitrium heterostichum snow-bed

Salix herbacea, Carex bigelowii, Juncus trifidus, Luzula spicata, Huperzia selago, Andreaea rupestris, Conostomum tetragonum, Pohlia nutans, Dicranum scoparium, Kiaeria starkei, Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Oligotrichum hercynicum, Polytrichastrum alpinum, Polytrichastrum sexangulare, Polytrichum piliferum, Barbilophozia floerkei, Diplophyllum albicans, Gymnomitrion concinnatum, Gymnomitrion corallioides, Lophozia sudetica, Marsupella brevissima, Nardia scalaris, Cetraria delisei, Cetraria islandica, Cladonia bellidiflora, Cladonia coccifera, Cladonia gracilis, Cladonia pyxidata, Cladonia uncialis, Cornicularia aculeata, Ochrolechia frigida, Gnaphalium supinum, Silene acaulis, Alchemilla alpina, Galium saxatile, Agrostis canina, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Nardus stricta